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Key: Scatopsidae
Key to European Scatopsidae, occasionally supplemented with some species from other regions.
References
Chandler, P.J., 1999. The British species of Anapausis Enderlein (Diptera, Scatopsidae). - Dipterists Digest 6: 1-21.
Freeman, P., & R.P. Lane, 1985. Bibionid and Scatopsid flies. - Handbooks for the Identification of British Insects 9(7): 1-74.
Haenni, J.-P., 1984. Contribution à la connaissance de la faune des Scatopsidae (Diptera) de Suisse. II. Le genre Anapausis Enderlein. - Mitteilungen der Schweizerischen Entomologischen Gesellschaft 57: 273-285.
Haenni, J.-P., 1997. Family Scatopsidae. In: Contributions to a manual of Palaearctic Diptera Volume 2 (L. Papp & B. Darvas, eds.): 255-272. Science Herald, Budapest.
Haenni, J.-P., 2013. A revision of the West Palaearctic species of Colobostema Enderlein, 1926 (Diptera, Scatopsidae). Part I. European subregion. - Mitteilungen der Schweizerischen Entomologischen Gesellschaft 86: 199-242.
Haenni, J.-P., &L. Pollini, 2015. Description of a new species of Parascatopse Cook from Switzerland, with taxonomic notes on other European species of the genus (Diptera, Scatopsidae). - Mitteilungen der Schweizerischen Entomologischen Gesellschaft 88: 65-72.
Krivosheina, N.P., 2000. A new and little known species of the genus Aspistes Meigen (Diptera, Scatopsidae) from Finland. - Entomologica Fennica 11: 119-123.
Last modified 25/11/2015 11:02 by Paul BeukNo of couplets: 37

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Key
1a
Wings normally developed.
2
b
Wings reduced to a minute vestigial lobe, halteres present. Whole surface of first abdominal tergite of male with a distinctive granulated ommatidia-like pattern.
Thripomorpha paludicola (Enderlein, 1905)
 
 
2a
Scutum with an elevated U-shaped ridge (Fig. 4). Anterior tibia with a strongly developed apical spur. Costa strongly swollen at junction with R4+5 (Fig. 8).
Aspistinae
3
b
Scutum without an elevated ridge, rounded or flattened in lateral view. Fore tibia not produced into an apical spur. Costa not swollen at junction with R4+5.
5
 
3a
Flagellum of antenna 5-segmented in both sexes. Fore femur simple, unarmed beneath (Holarctic, boreal).
Arthria analis (Kirby, 1837)
 
b
Flagellum of antenna 6-segmented in female, 8-10-segmented in male. Fore femur with spines beneath.
Three species in Fauna Europaea.
Aspistes Meigen, 1818
4
 
4a
Aspistes berolinensis Meigen, 1818
 
b
Aspistes helleni Krivosheina, 2000
 
 
5a
R4+5 and M fused for a distance, cross-vein R-M obliterated, stem of M fork arising from R4+5, well distally from transverse vein (base of R4+5) (Fig. 9). Hind tibia flattened, strongly swollen in apical third. A pseudotrachea on labella.
Four species in Fauna Europaea.
Ectaetiinae: Ectaetia Enderlein, 1912
 
b
Cross-vein R-M present, even if generally very short, often oblique or even horizontal, R4+5 and M separated, touching at most in one point (Figs 12-19) and base of M fork arising from just before or below transverse vein (base of R4+5). Hind tibia normal, at most slightly enlarged apically. Labella devoid of pseudotrachea.
6
 
6a
No false vein visible between M2 and CuA1 (Figs 10-11). Posterior veins and membrane with macrosetae among microtrichia. M fork incomplete, base of M1 lacking (Figs 10-11). Sperm pump of male directly attached to genital capsule.
Psectrosciarinae
7
b
False vein present as a concave fold between M2 and CuA1, sometimes bifurcated at apex (Figs 12-19) (this fold is not always very visible, the wing must be thoroughly observed under different angles of view; usually the fold appears clearly over a dark surface). Posterior veins and membrane with or without macrosetae among the microtrichia. M fork generally complete (except in Thripomorpha with well-visible false vein present). Sperm-pump of male attached to genitalia only by an elongated sperm-duct, lying apparently free in abdomen.
Scatopsinae
19
 
7a
Body strongly compressed laterally, elongated. R4+5 joining the costa gently in a smooth curve (Fig. 11).
Two species in Fauna Europaea.
Psectrosciara Kieffer in Enderlein, 1911
 
b
Body not strongly compressed laterally nor elongated. R4+5 joining rather abruptly the costa, forming with it an angle vicinous to 90 degrees (Fig. 10).
Twenty four species in Fauna Europaea, one described later from Central Europe by Haenni & Martinovský in 2014.
Anapausis Enderlein, 1912
8
 
8a
Vein M1 bent anteriorly at wing margin; macrotrichia on wing membrane anterior to M1. Male tergite 8 with two parallel strap-like curved arms, subapically joined by median bridge. Female sternite 7 with broad lateral lobes widely separated medially.
Anapausis talpae (Verrall, 1912)
 
b
Vein M1 not bent anteriorly at wing margin; macrotrichia mainly confined to membrane behind M1. Male tergite 8 without apically fused lateral arms. Female sternite 7 not divided into two lobes.
9
 
9a
First tarsomeres without comb-like rows of stout setae. Male tergite 8 rounded apically with small median indentation, lacking paired processes. Female tergite 8 with small projection medially; tergite 9 absent.
Anapausis nigripes (Zetterstedt, 1860)
 
b
First tarsomeres with comb-like rows of spiniform setae. Male tergite 8 with a pair of posteroventrally directed tapered blunt processes. Female tergite 8 without medial projection; tergite 9 well developed.
 
 
10a
Males.
11
b
Females.
15
 
11a
Sternite 6 broadly emarginate medially, lateral parts simple and not bilobed. Tergite 8 with pair of processes short, not longer than gap between them .
Anapausis rectinervis Duda, 1928
 
b
Sternite 6 deeply emarginate medially, lateral part not simple but variously lobed.
12
 
12a
Sternite 6 with lateral part not bilobed medially, but with apical margin angled internally and bearing a posteriorly directed thumb-like process near this angle. Tergite 8 with pair of processes about as long or a little longer than gap between them .
Anapausis pollicata Chandler, 1999
 
b
Sternite 6 with lateral part deeply bilobed medially. Tergite 8 with pair of processes longer than gap between them.
13
 
13a
Sternite 6 with narrow emargination, each lateral part with anterior margin evenly curved and posterior lobe pointed apically . Stemite 5 with large rounded anterior lobes, broader than the gap between them .
Anapausis dalmatina Duda, 1928
 
b
Sternite 6 with wider emargination, each lateral part with anterior margin concave on inner part and posterior lobe not pointed apically. Stemite 5 with rounded anterior lobes small and widely separated.
14
 
14a
Sternite 6 with posterior lobes only slightly indented subapically; tergite 8 dark brown, with processes shorter and more slender .
Anapausis soluta (Loew, 1846)
 
b
Sternite 6 with posterior lobes strongly indented subapically to give a hooked appearance; tergite 8 usually more reddish with processes longer and thicker .
Anapausis floricola Chandler, 1999
 
 
15a
Tergite 9 distinctly longer than broad, with straight basal margin and deeply inset in tergite 8 which is short medially. Stemite 8 emarginate apically.
Anapausis rectinervis Duda, 1928
 
b
Tergite 9 not longer than broad, with curved basal margin. Tergite 8 broader medially with curved apical margin.
16
 
16a
Stemite 9 produced apically to a point, protruding beyond tergite 9. Furca with a pair of diverging (ventrally directed) anterior processes (Fig. 9).
Anapausis sp. indet. 1
 
b
Stemite 9 rounded apically.
17
 
17a
Tergite 9 more angular and deeply inset in concave apical margin of tergite 8. Furca with single slender anterior process (Fig. 7).
Anapausis dalmatina Duda, 1928
 
b
Tergite 9 broadly ovate, more shallowly inset in tergite 8 (Figs 5-6).
18
 
18a
Tergite 8 without a rounded lobe laterally. Furca with a pair of short diverging (sometimes anteroventrally directed) anterior processes (Fig. 6).
Anapausis soluta (Loew, 1846)
 
b
Tergite 8 with bluntly rounded lateral lobe (extended more distally than in dalmalina). Furca with single elongate anterior process (Fig. 5).
Anapausis floricola Chandler, 1999
 
 
19a
Presence of macrosetae on posterior veins, and also on membrane in some genera (Figs 12-13). Stem of haltere without setae.
(In minute Parascatopse, at least 12 macrosetae present on CuA2, sometimes difficult to see; high magnification necessary).
Rhegmoclematini
20
b
No macrosetae on posterior veins and on membrane (Figs 14-19). Stem of haltere with setae in Palaearctic genera (except in extra-limital Lumpuria with quadrate scutum, dichoptic eyes in both sexes and elongated hind tibia).
25
 
20a
Presence of macrosetae on M1, M2, or both. M1 incomplete basally in Palaearctic species (Fig. 12). Flagellum of antenna 10-segmented. First abdominal tergite of male often with a granulated ommatidia-like structure.
Nine species in Fauna Europaea.
Thripomorpha Enderlein, 1905
 
b
No macrosetae on M1 and M2, macrosetae present only on CuA2 vein. M fork complete (Fig. 13).
21
 
21a
Flagellum of antenna 10-segmented.
Neorhegmoclemina Cook, 1955: Neorhegmoclemina catharinae Haenni, 1997
 
b
Flagellum of antenna 8-segmented.
22
 
22a
Male with sternite 7 shield-shaped . A regular distinct supraalar row of setae. Spiracular sclerite elongated, triangular.
Four species in Fauna Europaea.
Rhegmoclemina Enderlein, 1936
 
b
Male sternite 7 not shield-shaped. No distinct supraalar row of setae. Spiracular sclerite not elongated, rounded in general shape. Size very small, wing length about 1 mm (mostly coastal or halophilous).
Parascatopse Cook, 1955
23
 
23a
Legs with tibiae broadly whitish at base, tarsi and halteres pale brown.
Parascatopse minutissima (Verrall, 1886)
 
b
Legs uniformly darkish, though not as dark as body. Halteres at least darkened on knob.
24
 
24a
Sternite 7 broadly triangular. Lateral appendages in the genital capsule wide, rounded apically.
Parascatopse litorea (Edwards, 1925)
 
b
Sternite 7 with median posterior projection truncate apically (fig. 3). Lateral appendages in the genital capsule narrow, contorted.
Parascatopse distorta Haenni & Pollini, 2015
 
 
25a
Thorax stout, as wide as long, quadrate or nearly so. Wings densely microtrichiose.
Colobostematini
26
b
Thorax narrow, much longer than wide.
28
 
26a
A complete supernumerary cross-vein between M1 and R4+5 (cf. Fig. 15).
Four species in Fauna Europaea.
Holoplagia Enderlein, 1912
 
b
No complete supernumerary cross-vein between M1 and R4+5, but M1 often angled at 1/4 basal (Fig. 14), sometimes with a stump of vein directed towards R4+5 (Fig. 1).
27
 
27a
Flagellar segments with a distinct neck, short, bead-like or conical. Occiput regularly rounded posteriorly. Eyes dichoptic in female, hardly touching and nearly dichoptic in male. Thorax and abdomen dull, not shiny.
Eleven species in keyed by Haenni in 2013 but he mentions at least 18 Western Palaearctic species with probably more undescribed Mediterranean species.
Colobostema Enderlein, 1926
 
b
Flagellar segments devoid of neck, cylindrical and more or less elongated. Occiput hollowed out posteriorly, head short. Eyes clearly dichoptic in both sexes. Posterior tibia elongated.
Not yet recorded in the Palaearctic, 5 Oriental spp., Nepal; Haenni 1988b.
Lumpuria Edwards, 1928
 
 
28a
Wings infuscate, dull. A cluster of spiniform setae on sternite 7 posteriorly, beside usual setae (visible at high magnification). R4+5 long, joining costa at about 3/4 of wing length.
Two species in Fauna Europaea.
Ferneiella Cook in Freeman, 1985
 
b
Wings clear, more or less hyaline or milky, at most slightly infuscated. No spiniform setae on posterior margin of sternite 7. R4+5 of variable length.
29
 
29a
A complete supernumerary cross-vein between M1 and R4+5 (though translucent, this vein is
well-visible, especially under tangential light). Vein CuA2 strongly recurved towards wing base (Fig. 15). Tarsi white, contrasting with rest of body and legs.
Efcookella Haenni, 1998: Efcookella albitarsis (Zetterstedt, 1850)
 
b
No complete supernumerary cross-vein between M1 and M fork evenly curved or smoothly angled at 1/4 basal, with at most one anteriorly directed stump of vein arising towards R4+5.
30
 
30a
Second costal section practically equal to first section, or longer. R4+5 long, reaching costa at 2/3 of wing length or further (Figs 16-17); if hardly so (some Reichertella), then CuA2 smoothly curved towards wing tip (cf. Figs 16-17), not reaching abruptly hind margin of wing. Anterior spiracular sclerite nearly as high as long, with a comparatively large spiracular opening (Fig. 6). Palpus rounded, oval or truncate at apex, usually shorter or approximately as long as labella.
Scatopsini
31
b
Second costal section clearly shorter than first section. R4+5 short (Figs 18-19), reaching costa at middle of wing length or at most somewhat further, and then CuA2 reaching hind margin of wing abruptly, at a nearly right angle (Fig. 18). Palpus usually large, sausage shaped or pointed
at apex (Fig. 2), usually at least as long as labella or longer; anterior spiracular sclerite longer than high, spiracular opening comparatively smaller (Fig. 7). Thorax and abdomen entirely covered with a vestiture of minute microtrichia beside the usual pilosity (high magnification required).
Swammerdemellini
35
 
31a
Fork of M with an angle at 1/4-1/3 basal of M1 and an anteriorly directed stump of vein arising from this angle (translucent and sometimes weakly visible) (Fig. 1); if no stump of vein visible (Pharsoreichertella hamifera (Strobl)), then M fork strongly assymetrical, with M1 strongly bent and M2 nearly straight (Fig. 16).
32
b
Fork of M practically symmetrical, with M1 and M2 evenly and smoothly curved, without any indication of an angle or a stump of vein at 114-113 basal (Fig. 17).
34
 
32a
Tarsomere 1 of male hind leg shorter than tarsomere 2, or equal in length, armed with spines. Tergite 7 of male not projecting posteriorly, concealed under tergite 6 and weakly sclerotized. M1 with a well-marked angle at 1/4 basal and a stump of vein directed towards R4+5, well-visible though translucent (Fig. 1). Abdominal tergites and sternites bright, devoid of microtrichia beside the usual pilosity.
Three species in Fauna Europaea.
Scatopse Geoffroy, 1762
33
b
Tarsomere 1 of hind leg unmodified in male, longer than tarsomere 2. Tergite 7 of male well-visible and sclerotized, projecting posteriorly in 1 or 2 lobes. Angle of M1 not very marked, with a weakly visible anteriorly directed stump or, if stump absent, then fork of M strongly asymmetrical (Fig. 16). Abdominal tergites and sternites dull, at most subshiny, with a vestiture of microtrichia (visible at high magnification) beside the usual pilosity.
Three species in Fauna Europaea.
Pharsoreichertella Cook, 1974
 
 
33a
Entire length of vein R3 ventrally with 3-5 rows of sotulae; section 2 of C at least 1.25 times as long as section 3. Male: penis valves apically pointed or nearly so, claw-like (Fig. 10). Female: sternum 8 with a pair of valvifers.
Scatopse notata (Linnaeus, 1758)
 
b
Vein R3 ventrally with one row of setulae, at least on its proximal part; section 2 of C equal to section 3 or shorter. Male: penis valves apically broadly rounded. Female: sternum 8 without valvifers.
 
 
34a
Abdominal sternites (and also tergites) bright, devoid of microtrichia beside the usual pilosity. R4+5 long, reaching the costa in a smooth curve well beyond 2/3 of wing length (Fig. 17). Male pregenital segment (sternite and sometimes also tergite 7) diversely modified, with more or less developed, often asymmetrical lateral posterior projections. Penis elongated, filamentous, 2-3 branched, diversely modified at apex; 1-2 pairs of hypogynial valves in female.
Thirteen species in Fauna Europaea.
Apiloscatopse Cook, 1974
 
b
Abdominal sternites dull or at most subshiny, with a vestiture of microtrichia (visible at high magnification) beside the usual pilosity. R4+5 shorter, reaching the costa at about 2/3 of wing length, or if longer (Reichertella geniculata (Zetterstedt)), then running parallel to the costa and joining this vein after a marked knee. Male tergite 7 partly desclerotized or reduced, much shorter than sternite 7. Penis shorter and thick, more or less contorted, enlarged or flattened. No hypogynial valves in female.
Three species in Fauna Europaea.
Reichertella Enderlein, 1912
 
 
35a
M fork short or very short, always shorter than M stem . R1 and R4+5 joining costa very close to each other; distance between them (second costal section) shorter than length of R1 from origin of R4+5 to costa . Palpus pointed apically (Fig. 2).
Six species in Fauna Europaea, one species described later by Haenni from Spain in 2015.
Swammerdamella Enderlein, 1912
 
b
M fork obviously longer than stem (Fig. 18).
36
 
36a
Male genitalia laterally compressed. Palpus large, rounded.
Quateiella Cook, 1975: Quateiella inexpectata Haenni, 1988
 
b
Male genitalia not compressed laterally.
37
 
37a
Male tergite 7 asymmetrically produced posteriorly as a narrow spatulate process. Aedeagus elongated, coiled. Tarsi light coloured. Anterior spiracular sclerite a large, rather quadrate plate, twice as long as high.
Coboldia Melander, 1916: Coboldia fuscipes (Meigen, 1830)
 
b
Male tergite 7 symmetrical, produced or not posteriorly. Aedeagus short. Anterior spiracular sclerite triangular elongated (Fig. 7). Tarsi concolourous with rest of body and legs.
Nine species in Fauna Europaea.
Rhexoza Enderlein, 1936
 
 
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